| Citrate | Positive | Positive | Positive |
| Glucose | K/A Positive | A/A Positive | A/A Positive |
| Sucrose | Positive | Positive | Positive |
| Lactose | Positive | Positive | Positive |
| α-glucosidase | Positive | Negative | Negative |
| Urease | Positive | Negative | Negative |