| Characteristics | Results |
| Hydrolysis reaction | |
| Production of melanin pigment Nitrate reduction Gelatine reduction Coagulation of milk | negative positive positive negative |
| Carbon utilization of | |
| D-glucose D-xylose L-arabinose L-rhamnose D-fructose D-galactose raffinose D-mannitol Meso-inositol Salicin Sucrose | +++ +++ +++ +++ ++ +++ negative +++ ++ + + |