s2 = 2.76

s2 = 0.17

Fitted model Negative binomial

χ2 = 3.46

Fitted model Poisson

χ2 = 3.04

p < 0.05 df = 13

p < 0.01 df = 2

k = 0.07

External border

No nest

x ¯ = 0.12

s2 = 0.10

Fitted model Poisson

χ2 = 0.1

p < 0.01 df = 2

Second

No nest

x ¯ = 0.09

s2 = 0.08

Fitted model Poisson

χ2 = 0.05

p < 0.01 df = 2

Third

x ¯ = 1.14

x ¯ = 0.29

s2 = 8.68

s2 = 0.21

Fitted model Negative binomial

χ2 = 0.45

Fitted model Poisson

χ2 = 1.23

p < 0.05 df =13

p < 0.01 df = 2

k = 0.14

Fourth

x ¯ = 0.64

x ¯ = 0.34

s2 = 2.46

s2 = 0.23

Fitted model Negative binomial

χ2 = 0.02

Fitted model Poisson

χ2 = 1.56

p < 0.05 df = 5

p < 0.01 df = 2

k = 0.18

Center

x ¯ = 0.64

x ¯ = 0.26

s2 = 0.86

s2 = 0.21

Fitted model Negative binomial

χ2 = 0.0002

Fitted model Poisson

χ2 = 0.31

p < 0.05 df = 2

p < 0.01 df = 2

k = 2.23