| Variables | Description | Units | ||
| Map-based | Numerical | DTW | log10(DTW) | log10(m) |
| Slope | Slope (20 m focal average) | % | ||
| Canopy Closure | From forest inventory | % classes | ||
| Categorical | Aspect | Aspect (20 m focal average) | 4 Classes | |
| Ecosite | Bog, Fen, Marsh, Forest wetland, Shrub wetland, Riparian zone, Upland | 8 Classes | ||
| Forest Type | Forest Type (SW, HW, MX, Other) | 4 Classes | ||
| Plot-based | Numerical | L-Layer | Depth of litter layer | cm |
| VT | Vegetation Type (1-Poor to 4-Rich) | 4 Classes | ||
| Categorical | Microsite | Mound or pit | 2 Classes | |
| Soil Type | Organic vs. mineral soil | 2 Classes | ||