| Ego/boundary | attribute | Intergroup | Proactive aggression | Reactive aggression | Species (in gene without culture) |
| Territorial/rigid | Dualistic | ingroup- outgroup | high | low | human |
| Individualistic/ flexible | competitive-independent | extended outgroup | high | high | chimpanzee |
| Collectivistic/ flexible | cooperative-relational | extended ingroup | low | low | bonobo |