| Composition | Leaves | Needles |
| Lignin (mg∙g−1) | 301.40b | |
| Total carbohydrate (mg∙g−1) | 485.29b | |
| N (mg∙g−1) | | 7.99b |
| Lignin: N | 32.57b | |
| C: N | 52.44b | |