Habitat types

Number of burrows

Distance from river (m)

Distance from foot path (m)

Soil compaction (tones/cm ± SE)

Seashore

55

0.3

3.2

0.96 ± 0.13

Sand fill

30

2.6

0

0.26 ± 0.02

Footpath

23

7.0

1.2

2.04 ± 0.15