| Soil properties | Intact forest | Degraded forest | Significance | Agricultural land | Significance | Combined land uses | Significance |
| SOC (%) | 0.98 | 0.82 | *** | 0.83 | *** | 0.88 | NS |
| Total N (%) | 0.04 | 0.02 | *** | 0.03 | NS | 0.03 | NS |
| P (mg·kg−1) | 11.25 | 5.97 | * | 10.98 | NS | 9.40 | NS |
| CEC (cmol·kg−1) | 21.73 | 21.13 | NS | 17.05 | *** | 19.97 | NS |
| Ca2+ (cmol·kg−1) | 15.93 | 9.07 | *** | 12.14 | *** | 12.38 | ** |
| Mg2+ (cmol·kg−1) | 3.72 | 4.03 | NS | 3.31 | NS | 3.68 | NS |
| K+ (cmol·kg−1) | 0.97 | 0.67 | *** | 0.56 | *** | 0.73 | *** |
| Na+ (cmol·kg−1) | 0.19 | 0.11 | *** | 0.16 | *** | 0.15 | *** |
| pH in H2O | 5.96 | 5.99 | NS | 5.86 | NS | 5.94 | NS |